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One night out at sea, a mysterious Pokémon egg washes into a massive ship led by Phantom the Pirate. As Phantom is about to hatch the egg for his desires, one of his ship employees steps in, snatches the egg, and escapes the ship, being none other than a Pokémon ranger using his capture styler to make a clean getaway with a Mantine.

Lucario and the Mystery of Mew opens over the foggy landscape of Rota, with Ho-oh flying overhead. Then it goes to Lucario, who is jumping from rock to rock.

The story revolves around the Millennium Comet, which appears in the night sky for seven days once every thousand years. It is, at the same time, when the legendary Pokémon Jirachi awakens from a long slumber to absorb the comet's energy.

The film's setting is in the water-themed Johto city of Altomare (which means "High Sea"), based on Venice, Italy. The city is watched over by two legendary Pokémon, Latias and Latios.

The film focuses on Celebi who travels to the future and brings with a boy named Sam when being chased by a hunter. Meanwhile Ash, Misty, Brock, Pikachu and Togepi are on their way to the next gym in their Johto journey and get a lift from a man named Maki who takes them to his hometown of Arborville.

Professor Spencer Hale, once one of Professor Oak's top students and a good friend of the Ketchum family, is obsessed with discovering the secret behind the mysterious glyph-like Pokémon called Unown. One day while investigating some ruins, he disappears after encountering the enigmatic creatures, leaving his only daughter, Molly, alone and lonely back at his mansion.

Dr. Fuji, a research scientist of Team Rocket, briefly describes in a prologue how his researchers uncovered a fossil of the remnants of Mew, one of the rarest Pokémon in existence, believed to be extinct (although this is not the case).

A Pokémon Collector, Lawrence III, wishes to capture the three legendary birds and follow the legend's tale. This is all part of his plan to capture the Beast of the Sea, Lugia.

Shibusen, standing for Death Weapon Meister Academy (DWMA) the technical school for Reaper Armsmasters that operates from the deepest recesses of all the corners of the world, in order to prevent the second coming of the so-called Demon God who is feared upon bringing destruction of all humankind and envelop the world with nothing but madness and darkness. The reaper built this agency for weapons and for those who will be trained to wield them.

There once was a man named Gold Rodger, a pirate, and he was given the glorious title: ‘King of the Pirates’ for possessing the legendary treasure, One Piece. He had fame, fortune, and power, until one day, he caught and brought to the execution tower to be executed.

Ash is a young boy who has always dreamed of becoming a Pokemon trainer. At age 11 he is gifted his first Pokemon, Pikachu, from Professor Oak.