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Directed by Ootomo Katsuhiro of Memories and Steamboy. In the year 2019, thirty-one years after World War III, the authority of Neo-Tokyo is waging a constant battle against Underground rebels.

Gurren Lagann takes place in a fictional future where Earth is ruled by the Spiral King, Lordgenome, who forces mankind to live in isolated subterranean villages. These villages have no contact with the surface world or other villages, and are under constant threat of earthquakes.

A world controlled by a sinister brotherhood who hide their fangs behind masks of humanity, known as the Chronos Corporation. While investigating a mysterious explosion near his school, Sho Fukamachi happens upon the Chronos Corporation's Greatest weapon: a techno organic suit of bio armor known as "the Guyver.