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When an alien force eventually called the "Neuroi" attacks Earth around the 1940s, the world is devastated by destruction. However, one man is able to lead the invention of an anti-Neuroi weapon known as the "Striker unit" which augments magical power and also enables flight through two mechanical lower-leg coverings known as a sort of witches "broom.

The year is 1939 - it was that year that the Neuroi appeared. Nobody knows where they came from or what their ultimate agenda is, but the fact remains that their attacks drove people out of their towns and cities.

After discovering that Neuroi are capable of communication with humans and making peace with the nest above Gallia the Witches squads are now on a mission to restablish communication when a sudden attack vanishes the Neuroi, they now discover a new and massive nest that just appeared covering almost all Europe. Ruthless and without mercy it annihilates allied forces along with Witches.

In 1945, Yoshika Miyafuji, who lost her witch powers during the Strike Witches' last assignment, has been studying to become a doctor. Shizuka Hattori, one of her cadets in the Imperial Fuso Navy, then arrives to deliver a message: Yoshika is to be transferred for study abroad in Europe.

Ordinary life of high school students when in all honesty it is nothing but normal. theirs a kindergarten genius who built her own mom with a giant gear in her back and cake in her limbs and high school students who have some serious problems and a talking cat.

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