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Rintaro Okabe is a self-proclaimed "mad scientist" who believes that an international scientific organization named SERN is conspiring to reshape the world according to its own interests. He and his friend Itaru Hashida inadvertently create a gadget able to send messages to the past.

Koyomi Araragi is a student who has recently survived a vampire attack, with the help of his friend Hanekawa Tsubasa. Now, he finds that she has been possessed by an oddity called the Curse Cat, and is now attacking people, including her parents.

Gurren Lagann takes place in a fictional future where Earth is ruled by the Spiral King, Lordgenome, who forces mankind to live in isolated subterranean villages. These villages have no contact with the surface world or other villages, and are under constant threat of earthquakes.