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The end of the Neon Genesis Evangelion left many questions as to the fate of NERV, the Children, the Angels, and many other plot factors unanswered, unexplained, or otherwise hanging. To answer a legion of now befuddled Evangelion fans, Gainax remade the ending into a two-part movie known as Death and Rebirth which was to supersede episodes 25 and 26 of the television series, which then led to this movie, Death and Rebirth itself being recreated into part one of The End of Evangelion.

In 2015 huge, tremendously powerful alien war machines return to Tokyo. The only hope for Mankind's survival lies in the Evangelion, a humanoid fighting machine developed by NERV, a special United Nations agency.

While fighting to save his village from the attack of a demon-god Ashitaka comes in contact with it and is inflicted with a deadly curse. In his search for a cure he is forced to leave his village forever and journey to the forest inhabited by animal gods.