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Aster and Yuno were abandoned together at the same church, and have been inseparable since. As children, they promised that they would compete against each other to see who would become the next sorcerous emperor.

The story revolves around Rin Okumura, a teenager possessing abnormal strength and endurance who, along with his younger twin brother Yukio, was raised by Father Shiro Fujimoto. Soon after Yukio departs to study at the prestigious True Cross Academy, Rin learns that he is the son of Satan, the strongest of all demons.

In the year 2022, the gaming industry has reached a new height after the development of the NerveGear, a helmet-like controller that allows the user to fully dive their senses into the world of virtual reality. After a closed beta with only 1,000 testers, the Virtual Reality MMORPG Sword Art Online is launched.

Set in a Japan after the apocalypse in the future in 2071, the world has been almost entirely destroyed by these mysterious monsters known as the Aragami. An organization calling itself Fenrir was established to exterminate Aragami using 'God Arcs', weapons specially made from Aragami cells.

Gurren Lagann takes place in a fictional future where Earth is ruled by the Spiral King, Lordgenome, who forces mankind to live in isolated subterranean villages. These villages have no contact with the surface world or other villages, and are under constant threat of earthquakes.