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Equipped with technologically advanced weaponry and specialized elite troops, private military organization Mithril covertly protects the Earth’s peace and order from the threat of destruction and terrorism. The backbone of Mithril’s operations rely on the Whispered, a group of exemplary and intelligent individuals, responsible for the invention and production of powerful machinery.

Gurren Lagann takes place in a fictional future where Earth is ruled by the Spiral King, Lordgenome, who forces mankind to live in isolated subterranean villages. These villages have no contact with the surface world or other villages, and are under constant threat of earthquakes.