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Tanaka Yukio, better known by his nickname Koyuki is a 14-year-old who feels disconnected from life itself. Through the act of saving a mismatched dog, he meets guitarist Minami Ryuusuke and becomes involved in Ryuusuke's new band BECK.

With dreams of becoming a composer and someday writing a song for her favorite idol, Haruka enters the Saotome Academy, a prestigious performing arts school. If successful there, she’ll be able to join the Shining Agency after graduation.

One evening in fall, with one month left before the school festival, the last member of the ruined light music club, Haruki Kitahara, looks out the window and plays his guitar in the direction of the festival stage set up outside. This was the simple little adventure that the conscientious honor student had hoped for two and a half years to achieve before his graduation in half a year.

The story is set in the Bakumatsu era, at the end of the shoguns' rule over Japan in the middle of the 19th century. The Tokugawa shogunate uses the brainwashing Heaven's Songs by the top idols in Shinsengumi to subjugate the country and its people.

Every day, a young girl wearing a mask stands by the beach and sings a nostalgic melody. After experiencing two sudden heart-wrenching partings when she was only a child, Nino Arisugawa has been singing her songs to the ocean, bound by a promise made with her two childhood friends—her first love, Momo Sakaki, and a boy who composed music, Kanade "Yuzu" Yuzuriha.

The music anime will follow the daily lives and associated drama of four of TsukiPro's very unique groups — SolidS, Quell, SOARA, and Growth