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The story takes place in Seikyou Private School. A facility that consists of multiple buildings that connect together to accommodate both junior and senior high school students.

Mutated homo sapiens that have a dangerous telekinetic power is being captured and isolated in laboratories by the government. One of the sixth-sense-possessing humanoids escapes loses her memories and winds up living with a boy from her past.

The film's setting is in the water-themed Johto city of Altomare (which means "High Sea"), based on Venice, Italy. The city is watched over by two legendary Pokémon, Latias and Latios.

The film focuses on Celebi who travels to the future and brings with a boy named Sam when being chased by a hunter. Meanwhile Ash, Misty, Brock, Pikachu and Togepi are on their way to the next gym in their Johto journey and get a lift from a man named Maki who takes them to his hometown of Arborville.

Chizuru Yukimura heads to Kyoto to search for her father. While there, she witnesses a fight between Rasetsu & members of the Shinsengumi.

There are many idols with long-established talent agency 346 Production. And now the company is starting a new program, the Cinderella Project!